Joined: Nov. 2006
You have never heard about Oudemanse and after him called phenomenon of the total coloration of animals ("Totalzeichnung"). That's why Alan pursued me - instead of making his own research - to ISCID with his questions. †
Because you didn't find any information using google the phenomenon apparently does not exist for you. †
But you should learn more about how to use google:
The independence of pattern formation mechanisms means that the coordination of united patterns of fore- and hindwings is accidental. This is remarkable, because from Oudemansís principle  , patterns appearing on the exposed surface of fore- and hindwing at the natural resting position are often integrated to form a composite and united adaptive pattern with their surrounding environment.
VMartin can't understand how a particular butterfly evolved to look the way it does, therefore goddidit.
But we are pretty sure naturalselectiondidnotdoit.
I could not answer, but should maintain my ground.-