Joined: Nov. 2006
This is the picture of the Issoria lathonia, where the reverse side of the butterfly wings can be observed:
The color patterns of front wings and hind wings make perfect harmony. But only in these resting position of wings, when the front wings are behind hind wings. We see only small parts of reverse side of front wings. The hided, covered part of them have almost the same coloration as front parts of butterfly wings coloration.
Do you see the white spots on the upper margin at the front wings? There are many black spots on wings which look like copy of spots on the upper part of wings.
So the harmony arise only in special position of wings. It seems like an artist painted the pattern in this exactly position. The phenomenon - neglected nowadays - is called after entomologist Oudemanse (or die Totalzeichnung, the term coined by Suffert) .
We often see animal coloration making the whole nice picture. This picture "is painted" over many parts of animal body which developes independently during ontogenesis.
Heikertinger considered "natural selection" as totally wrong explanatin of the phenomenon conceived in heads of "Hypothetiker" as he called proponents of natural selection.
I couldn't find a picture of Papilio dolicaon where he made his issue about the Oudemanse effect. There are semi-circles on its front and hind wings that fit into each other and create circles in the resting position of wings. It is hardly imaginable (if you are not a darwinist of course) that predators left only those individuals where circles were perfect and eliminated all those individuals, which didn't create perfect circles. It would mean that predators have also some aesthetical feeling.
I could not answer, but should maintain my ground.-